(1Shandong University of Science and Technology, Qingdao 266590, Shandong, China;2CAS Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China; 3Huitong Experimental Station of Forest Ecology of Chinese Academy of Sciences, Shenyang 110016, China; 4Qingyuan Forest CERN, Chinese Academy of Sciences, Shenyang 113300, China).