Abstract: Species competition is one of the important ecological courses that influence the evolvement of ecosystem. To an ecosystem of predator-prey, competition is always bidirectional, that is there is competitive infection on superior species and inferior species. Without regarding to the destruction of habitat, the bidirectional competitive mechanism was introduced, Tilman’ unilateral competitive model was extended to bidirectional competitive model of metapopulation. At the same time, the competitive dynamics of the 6-metapopulation species are simulated on computer. The results showed that at the equilibrium, the coexistence condition of population competition is migratory ability and competition ability take on exponential negative correlation, the outcome of competition makes the superior and inferior sequence changed, the exclusion and coexistence of species competition was affected by migratory ability and competition ability, the metapopulations which are excluded in local patches can exist in regional scale, that is escaping coexistence.

Key words: Environmental factors, Recrudescence of gonad, Ovulation, Spawning