[1] Li HR, Reynolds JF. On definition and quantification of heterogeneity. Oikos, 1995, 73: 280-284 [2] 高梅香, 刘冬, 张雪萍, 等. 三江平原农田地表和地下土壤螨类丰富度与环境因子的空间关联性. 生态学报, 2016, 36(6): 1782-1792 [Gao M-X, Liu D, Zhang X-P, et al. Spatial relationships between the abundance of aboveground and belowground soil mite communities, and environmental factors in a farmland on the Sanjiang Plain, China. Acta Ecologica Sinica, 2016, 36(6): 1782-1792] [3] Webster R, Boag B. Geostatistical analysis of cyst nematodes in soil. Journal of Soil Science, 1992, 43: 583-595 [4] Gutiérrez-López M, Jesús JB, Trigo D, et al. Relationships among spatial distribution of soil microarthropods, earthworm species and soil properties. Pedobiologia, 2010, 53: 381-389 [5] Hortal J, Roura-Pascual N, Sanders NJ, et al. Understanding (insect) species distributions across spatial scales. Ecography, 2010, 33: 51-53 [6] Heino J, Alahuhta J. Elements of regional beetle faunas: Faunal variation and compositional breakpoints along climate, land cover and geographical gradients. Journal of Animal Ecology, 2015, 84: 427-441 [7] Tonkin JD, Stoll S, Jähnig SC, et al. Contrasting metacommunity structure and beta diversity in an aquatic-flood-plain system. Oikos, 2016, 125: 686-697 [8] Guo Y, Gao MX, Liu J, et al. Disentangling the dri-vers of ground-dwelling macro-arthropod metacommunity structure at two different spatial scales. Soil Biology & Biochemistry, 2019, 130: 55-62 [9] 朱纪元, 李景科, 高梅香, 等. 帽儿山红松人工林鞘翅目成虫群落小尺度空间异质性变化特征. 生态学报, 2017, 37(6): 1975-1986 [Zhu J-Y, Li J-K, Gao M-X, et al. Spatially heterogeneous dynamics of adult Coleoptera communities at a small scale in a Pinus koraiensis plantation on Maoer Mountain. Acta Ecologica Sinica, 2017, 37(6): 1975-1986] [10] 胡媛媛, 朱纪元, 闫龙, 等. 温带落叶阔叶林地表鞘翅目成虫小尺度空间格局动态分析. 生态学报, 2018, 38(5): 1841-1851 [Hu Y-Y, Zhu J-Y, Yan L, et al. Analysis of the dynamic spatial pattern of adult Coleoptera communities at fine scale in a temperate deci-duous broad-leaved forest. Acta Ecologica Sinica, 2018, 38(5): 1841-1851] [11] 倪娟平, 程赛赛, 高梅香, 等. 丰林典型阔叶红松林地表鞘翅目成虫空间异质性及其与环境因子的空间关联性. 生物多样性, 2018, 26(1): 14-26 [Ni J-P, Cheng S-S, Gao M-X, et al. Spatial heterogeneities of ground-dwelling Coleoptera adults and their spatial correlations with environmental factors in a typical broad-leaved Korean pine forest in the Fenglin Nature Reserve. Biodiversity Science, 2018, 26(1): 14-26] [12] 刘霞, 赵东, 程建伟, 等. 放牧和刈割对内蒙古典型草原大型土壤动物的影响. 应用生态学报, 2017, 28(6): 1869-1878 [Liu X, Zhao D, Cheng J-W, et al. Effects of grazing and mowing on macrofauna communities in a typical steppe of Inner Mongolia, China. Chinese Journal of Applied Ecology, 2017, 28(6): 1869-1878] [13] Gongalsky KB, Cividanes FJ. Distribution of carabid beetles in agroecosystems across spatial scales: A review. Baltic Journal of Coleopterology, 2008, 8: 15-30 [14] 任国栋, 于有志. 中国荒漠半荒漠的拟步甲科昆虫. 保定: 河北大学出版社, 1999 [Ren G-D, Yu Y-Z. Tenebrionidae Fauna in Desert and Semi-Desert Areas in China. Baoding: Hebei University Press, 1999] [15] Ayal Y. Trophic structure and the role of predation in shaping hot desert communities. Journal of Arid Environments, 2007, 68: 171-187 [16] 梁存柱, 朱宗元, 李志刚. 贺兰山植被. 银川: 阳光出版社, 2012 [Liang C-Z, Zhu Z-Y, Li Z-G. The Vege-tation of Helan Mountain. Yinchuan: Sunshine Press, 2012] [17] 杨贵军, 于有志, 王新谱. 宁夏贺兰山拟步甲科的区系组成与生态分布. 宁夏大学学报: 自然科学版, 2011, 32(1): 67-72 [Yang G-J, Yu Y-Z, Wang X-P. Tenebrionidae fauna and ecological distribution in Ningxia Helan Mountain Nature Reserve. Journal of Ningxia University: Natural Science, 2011, 32(1): 67-72] [18] 杨贵军, 王敏, 杨益春, 等. 贺兰山甲虫物种丰富度分布格局及其环境解释. 生物多样性, 2019, 27(12): 1309-1319 [Yang G-J, Wang M, Yang Y-C, et al. Distribution patterns and environmental interpretation of beetle species richness in Helan Mountain of northern China. Biodiversity Science, 2019, 27(12): 1309-1319] [19] 莫多闻, 朱忠礼, 万林义. 贺兰山东麓冲积扇发育特征. 北京大学学报: 自然科学版, 1999, 35(6): 816-823 [Mo D-W, Zhu Z-L, Wan L-Y. The alluvial fans along the eastern foot of Helan Mountain. Acta Scientiarum Naturalium Universitatis Pekinensis, 1999, 35(6): 816-823] [20] 杨益春, 杨贵军, 王杰. 地形对贺兰山步甲群落物种多样性分布格局的影响. 昆虫学报, 2017, 60(9): 1060-1073 [Yang Y-C, Yang G-J, Wang J. Effects of topographic factors on the distribution pattern of carabid species diversity in the Helan Mountain, northwestern China. Acta Entomologica Sinica, 2017, 60(9): 1060-1073] [21] 赵亚楠, 周玉蓉, 王红梅. 宁夏东部荒漠草原灌丛引入下土壤水分空间异质性. 应用生态学报, 2018, 29(11): 3577-3586 [Zhao Y-N, Zhou Y-R, Wang H-M. Spatial heterogeneity of soil water content under introduced shrub (Caragana korshinskii) in desert grassland of the eastern Ningxia, China. Chinese Journal of Applied Ecology, 2018, 29(11): 3577-3586] [22] 马克平, 刘玉明. 生物多样性测度方法: α-多样性. 生物多样性, 1994, 2(4): 231-239 [Ma K-P, Liu Y-M. Measurement of biotic community diversity. I: α diversity (Part 2). Biodiversity Science, 1994, 2(4): 231-239] [23] 王政权. 地统计学及在生态学中的应用. 北京: 科学出版社, 1999 [Wang Z-Q. Geostatistics and Its Application in Ecology. Beijing: Science Press, 1999] [24] Jung WK, Kitchen NR, Sudduth KA, et al. Spatial characteristics of Claypan soil properties in an agricultural field. Soil Science Society of America Journal, 2006, 70: 1387-1397 [25] 王正军, 李典谟, 商晗武, 等. 地质统计学理论与方法及其在昆虫生态学中的应用. 昆虫知识, 2002, 39(6): 405-411 [Wang Z-J, Li D-M, Shang H-W, et al. Theories and methods of geostatistics and its their application in insect ecology. Entomological Knowledge, 2002, 39(6): 405-411] [26] Cambardella CA, Moorman TB, Karlen DL, et al. Field-scale variability of soil properties in central Iowa soils. Soil Science Society of America Journal, 1994, 58: 1501-1511 [27] 高梅香, 何萍, 刘冬, 等. 温带落叶阔叶林土壤螨群落多尺度空间自相关性. 土壤通报, 2014, 45(5): 1104-1112 [Gao M-X, He P, Liu D, et al. Multi-scale spatial autocorrelation of soil mite community in a temperate deciduous broad-leaved forest, Northeast China. Chinese Journal of Soil Science, 2014, 45(5): 1104-1112] [28] 史文娇, 岳天祥, 石晓丽, 等. 土壤连续属性空间插值方法及其精度的研究进展. 自然资源学报, 2012, 27(1): 163-175 [Shi W-J, Yue T-X, Shi X-L, et al. Research progress in soil property interpolators and their accuracy. Journal of Natural Resources, 2012, 27(1): 163-175] [29] Santos ADL, Gómez-González LA, Alonso C, et al. Adaptive trends of darkling beetles (Col. Tenebrionidae) on environmental gradients on the island of Tenerife (Canary Islands). Journal of Arid Environments, 2000, 45: 85-98 [30] Parmenter RR, Parmenter CA, Cheney CD. Factors influencing microhabitat partitioning in arid-land darkling beetles (Tenebrionidae): Temperature and water conservation. Journal of Arid Environments, 1989, 17: 57-67 [31] Mazía CN, Chaneton EJ, Kitzberger T. Small-scale habitat use and assemblage structure of ground-dwelling beetles in a Patagonian shrub steppe. Journal of Arid Environments, 2006, 67: 177-194 [32] Bartholomew A, Hafezi SA, Karimi S. Effects of habitat complexity on the abundance, species richness and size of darkling beetles (Tenebrionidae) in artificial vegetation. Journal of Arid Environments, 2016, 129: 35-41 [33] Bartholomew A, Moghrabi JE. Seasonal preference of darkling beetles (Tenebrionidae) for shrub vegetation due to high temperatures, not predation or food availability. Journal of Arid Environments, 2018, 156: 34-40 [34] Gao MX, Qiao ZH, Hou HY, et al. Factors that affect the assembly of ground-dwelling beetles at small scales in primary mixed broadleaved-Korean pine forests in north-east China. Soil Ecology Letters, 2020, 2: 47-60 |